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Yen, Shen-Horn



Yen, Shen- Horn


Associate Professor


Systematic Biology
Mimicry Biology
Plant/Insect Relationships
Biodiversity Database
Aquatic Botany



886-7-5252000 Ext:3612



Highest Education: Imperial College London , Ph.D. (2004)

Research Interests

Systematics, phylogenetics and evolutionary ecology of the Lepidoptera and some smaller insect orders


Evolutionary dynamics and behavioural ecology relevant to mimicry, camouflage and aposematism


Colonization and diversification of phytophagous insects on plants


Technology and applications of biodiversity and natural history collection databases

Papers in peer reviewed journals


Prathapan KD, Pethiyagoda R, Raven PH...Yen SH...Zhou HZ (2018) When the cure kills - CBD limits biodiversity research. Science 360(6396): 1405.


Wang, MY, Vasa V, Chittka L, Yen SH (2017) Sheep in wolf's clothing: multicomponent traits enhance the success of mimicry in spider mimicking moths. Animal Behaviour, 127: 219-224.
3 Wei CH, Yen SH (2017) Mimaporia, a new genus of Epicopeiidae (Lepidoptera), with description of a new species from Vietnam. Zootaxa, 4254: 537-550.
4 Wei CH, Lohman DJ, Peggie, D, Yen SH (2017) An illustrated checklist of the genus Elymnias Hubner, 1818 (Nymphalidae, Satyrinae). ZooKeys, 676: 47-152.
5 Wu LW, Yen SH, Lees DC, Lu CC, Yang PS, Hsu YF (2015) Phylogeny and Historical Biogeography of Asian Pterourus Butterflies (Lepidoptera: Papilionidae): A Case of Intercontinental Dispersal from North America to East Asia. PLoS One, 10(10):e0140933.
6 Regier JC, Mitter C, Kristensen NP, Davis DR, Nieukerken EJ van, Rota J, Simonsen TJ, Mitter KT, Kawahara AY, Yen SH, Cummings MP, Zwick A (2014) A molecular phylogeny for the oldest (non-ditrysian) lineages of extant Lepidoptera, with implications for classification, comparative morphology and life history evolution. Systematic Entomology, 40(4): 671-704
7 Yu F, Chen FQ, Yen SH, Tu LH, Zho CD, Guerrieri E, Zhang YZ (2014) Preliminary phylogeny of the genus Copidosoma (Hymenoptera, Encyrtidae), polyemrbyonic parasitoids of Lepidoptera.Systematic Entomology, 39(2): 325-334.
8 Tamaska T, Kertesz K, Veretesy Z, Balint Zs, Kun A, Yen SH, Biro LP. (2013) Colour changes upon cooling of Lepidoptera scales containing photonic nanoarchitectures. Journal of Insect Science, 13: 87
9 Wu, S., Owada M, Hsiao SF, Yen SH (2013) Review of the genus Cidariplura Butler, 1879 (Lepidoptera, Erebidae, Herminiinae) in Taiwan with descriptions of four new species. Zootaxa, 3746 (1): 143-160.
10 Nieukerken, EJ van, Kaila L, Kitching IJ, Kristensen NP, Lees DC, Minet J, Mitter C, Mutanen M, Regier JC, Simonsen TJ, Wahlberg N, Yen SH, Zahiri R, Adamski D, Baixeras J, Bartsch D, Bengtsoon BA, Brown JW, BUcheli SR, Davis DR, de Prins J, de Prins W, Epstein ME, Gentili-Poole P, Gielis C, Hattenschwiler P, Hausmann A, Holloway JD, Kallies A, Karsholt O, Kawahara AY, Koster S, Kozlov MV, Lafontaine JD, Lamas G, Landry J-F, Lee S, Nuss M, Park K-T, Penz C, Rota J, Schintlmeister A, Schmidt BC, Sohn J-C, Solis MA, Tarmann GM, Warren AD, Weller S, Yakovlev RV, Zolotuhin V, Zwick A (2011) Order Lepidoptera Linnaeus, 1758. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 212-221.
11 Wu LW, Lees DC, Yen SH, Lu CC, Hsu YF (2010)[2011] The complete mitochondrial genome of the near-threatened swallowtail, Agehana maraho (Lepidoptera: Papilionidae): evaluating sequence variability and suitable markers for conservation genetic studies. Entomological News 121(3): 267-280.(publishing date postponded until 2011)
12 Wu LW, Yen SH, Lees DC, Hsu YF (2010) Elucidating genetic signatures of native and introduced populations of the Cycad Blue, Chilades pandava to Taiwan: a threat both to Sago Palm and to native Cycas populations worldwide. Biological Invasions 12(8): 2649-2669.
13 Scott JL, Kawahara AY, Skevington JH, Yen SH, Sami A, Smith ML, Yack JE (2009) Talking evolves from walking in terrestrial caterpillars. Nature Communications Article number 4 doi: 10.1038/ncomms1002.
14 Wu LW, Yen SH, YF Hsu (2009) An introduced plant induced a native butterfly outbreak that may threatean the survival of an endangered plant. Insect and Nature 44(13): 14-17.
15 Regier JC, Zwick A, Cummings MP, Kawahara AK, Cho S, Weller S, Roe A, Baixeras J, Brown JW, Parr C, Davis DR, Epstein M, Hallwachs W, Hausmann A, Janzen DH, Kitching IJ, Solis A, Yen SH, Bazinet AL, Mitter C (2009) Toward reconstructing the evolution of advanced moths and butterflies (Lepidoptera: Ditrysia): an initial molecular study. BMC Evolutionary Biology 2009(9): 280.
16 Vila M, Mari-Mena N, Yen SH, Lopez-Vaamonde C (2009) Characterization of ten polymorphic microsatellite markers for the protected Spanish moon moth Graellsia isabelae (Lepidoptera: Saturniidae). Conservation Genetics DOI: 10.1007/s10592-009-9905-1
17 Wei CH, Kun A, Yen SH (2007) A preliminary analysis of the phylogeny of the Ethminae moths (Lepidopteran: Gelechioidea) with special reference to the evolutionary pattern of host use. Acta Zoological Academiae Sientiarum Hungaricae 53 (Suppl.1): 61-100.
18 Yen SH, Minet J (2007) Cimelioidea: a new superfamily name for the gold moths (Lepidoptera: Glossata). Zoological Studies 46 (3): 262-271.
19 Hsu YF, Yen SH (2006) Immature biology of Prosotas reveals an oviposting strategy unusual in polyommatine lycaenids (Lepidoptera, Lycaenidae, Polyommatinae), with description of a new subspecies of P. dubiosa from Taiwan. Zoological Studies 45: 308-321.
20 Faucheux MF, Kristensen NP, Yen SH (2006) The antennae of neopseustid moths: morphology and phylogenetic implications, with special reference to the sensilla (Insecta, Lepidoptera). Zoologischer Anzeiger 245: 131-142.
21 Hsu YF, Chiba H, Yen SH (2006) Celaenorrhinus ratna Fruhstorfer, 1908 (Insecta, Lepidoptera): proposed conservation of the specific name. Bulletin of Zoological Nomenclature 63(1): 114-117.
22 Niehuis O, Yen SH, Naumann CM, Misof B (2006) Higher phylogeny of zygaenid moths (Insecta: Lepidoptera) inferred from nuclear and mitochondrial sequence data and the evolution of larval cuticular cavities for chemical defence. Molecular Phylogenetics and Evolution 39: 812-829.
23 Yen SH, Robinson GS, Quicke DLJ (2005) Phylogeny, systematics and evolution of mimetic wing patterns of Eterusia moths (Lepidoptera: Zygaenidae: Chalcosiinae). Systematic Entomology 30:358-397.
24 Yen SH (2005) Case 3295. Eterusia cingala Moore, 1877 (Insecta, Lepidoptera): proposed precedence over Heterusia aedea var. septentrionicola C. Felder et R. Felder, 1862. The Bulletin of Zoological Nomenclature 62(1): 18-20.
25 Sohn JC, Yen SH (2005) A taxonomic revision on Epipleminae (Lepidoptera: Uraniidae) of Korean Peninsula. Zoological Studies 44: 44-70.
26 Yen SH, Robinson GS, Quicke DLJ (2005) The Phylogenetic relationships of Chalcosiinae (Lepidoptera, Zygaenoidea, Zygaenidae). Zoological Journal of the Linnean Society 143: 161-341.
27 Yen SH, Solis MA, Goolsby J (2004) Austromusotima, a new musotimine genus (Lepidoptera: Crambidae) feeding on old world climbing fern, Lygodium microphyllum (Schizaeaceae). Annals of the Entomological Society of America 97: 397-410.
28 Yen SH (2004) The nomenclatural and systematic problems surrounding Eterusia aedea (Linnaeus, 1763) (Lepidoptera, Zygaenidae, Chalcosiinae). Zoological Studies 43: 20-34.
29 Solis MA, Yen SH, Goolsby J (2004) Musotimines from Southeast Asia Feeding on Lygodium microphyllum, Lygomusotima stria, New Genus and Species, and Redescription of Neomusotima conspurcatalis Warren, New Combination (Lepidoptera: Crambidae). Annals of the Entomological Society of America 97: 64-76.
30 Quicke DLJ, Yen, SH, Shaw MR (2004) First host record for the rogadine genus Conspinaria (Hymenoptera: Braconidae), and notes on rogadine and zygaenid parasitoid associations. Journal of Natural History 38: 1437-1442.
31 Hsu YF, Ding DS, Yen SH, Qian ZQ (2004) Systematic problems surrounding Howarthia melli (Forster) (Lepidoptera, Lycaenidae, Theclinae), with description of a new species and a review of the Rhododendron-association in lycaenid butterflies. Annals of the Entomological Society of America 97: 653-666.
32 Als T, Vila R, Kandul N, Nash D, Yen SH, Hsu YF, Mignault A, Boomsma K, Pierce (2004) The evolution of alternative parasitic life histories in Large Blue butterflies. Nature 432: 286-290.
33 Quicke DLJ, Yen SH, Shaw MR (2004)[2003] First host record for the rogadine genus Conspinaria (Hymenoptera: Braconidae), and notes on rogadine and zygaenid parasitoid associations. Journal of Natural History 38: 1437-1442.
34 Yen SH, Kitching IJ, Tzen CS (2003) A new subspecies of hawkmoth from Lanyu, Taiwan, with a revised and annotated checklist of the Taiwanese Sphingidae (Lepidoptera). Zoological Studies 42 (2): 292-306.
35 Yen SH (2003) A new species of Soritia from Taiwan, with phylogenetic comments on the genus. Nachrichten des Entomologischen Vereins Apollo, Frankfurt am Main, N.F. 24(1/2): 39-46.
36 Yen SH (2003) A new species of Eterusia from Mindanao and Seram (Lepidoptera, Zygaenidae, Chalcosiinae). Entomologisches Zeitschrift 126: 121-126.
37 Yen SH, Nassig WA, Naumann S, Brechlin R (2000) A new species of the miranda-group of the genus Loepa from Taiwan (Lepidoptera: Saturniidae). Nachrichten des Entomologischen Vereins Apollo 21 (3): 153-162.
38 Fanger H, Yen SH, Naumann CM (1999) External morphology of the last instar larva of Phauda mimica Strand, 1915 (Lepidoptera: Zygaenoidea). Entomologica scandinavica 29: 429-450.
39 Hsu YF, Yen SH (1998) Notes on Boloria pales yangi ssp.nov., a remarkable disjunction in butterfly biogeography (Lepidoptera: Nymphalidae). Journal of Researches on Lepidoptera 34: 142-146.
40 Yen SH, Yang PS (1998) The genus Soritia of Taiwan (Zygaenidae, Chalcosiinae, Chalcosiini). Tinea 15 (3): 207-220.
41 Yen SH, Yang PS (1997) Two new genera of Chalcosiinae (Zygaenidae) from eastern Palaearctic Asia. Transactions of The Lepidopterological Society of Japan 48 (4): 243-263.
42 Yang YP, Yen SH (1997) Notes on Limnophila (Scrophulariaceae) of Taiwan. Botanical Bulletin of Academia Sinica (1997) 38: 285-295.
43 Yen SH, Kishida Y (1997) Revision on Herimba atkinsoni Moore, 1879 (Thyrididae: Striglininae) and descriptions of three new subspecies from China and Taiwan. Transactions of the Lepidopterological Society of Japan 48 (2): 97-108.
44 Yen SH (1997) Pryeria sinica Moore (Lepidoptera, Zygaenidae), a newly discovered relic in Taiwan. Transactions of the Lepidopterological Society of Japan 48 (1): 39-48.
45 Yen SH (1997) Scaptesyle bicolor integra Swinhoe, a new addition to Lithosiinae (Arctiidae) of Taiwan. Tinea 15 (2): 94-100.
46 Yen SH (1997) Pseudeuchromia maculifera catachroma Schultza, 1907 (Geometridae, Ennominae), a new addition to the moth fauna of Taiwan. Transactions of the Lepidopterological Society of Japan 48 (1): 18-24.
47 Yen SH, Yang PS (1996) Uthinia albistrigbalis (Hampson) (Pyralidae: Scopariinae) from Taiwan. Japanese Heterocerists’ Journal 187: 189-191.
48 Yen SH, Jean JL, Yang PS (1996) On two Artona species from Taiwan (Lepidoptera: Zygaenidae, Procridinae). Entomologist' Gazette 47: 99-108
49 Yen SH (1996) Aeolopetra lanyuensis Yen, a new pyralid moth (Pyralidae, Musotiminae) from Taiwan. Tinea 14 (4): 293-299.
50 Yen SH (1996) A new addition to Campylotes (Lepidoptera, Zygaenidae, Chalcosiinae) of Taiwan. Tinea 14 (4): 289-292.
51 Yen SH, Yu HS, Mu JH, Tan HJ (1995) On Lyssa zampa (Butler, 1867) (Uraniidae) from Taiwan. Japanese Heterocerists’ Journal 186: 173-175.
52 Yen SH, Mu JH, Jean JL (1995) The life histories and biology of Epicopeiidae of Taiwan. Transactions of the Lepidopterological Society of Japan 36 (4): 175-184.
53 Yen SH, Jean JL, Su JP (1995) On the first record of Tirumala limniace orestilla Fruhstorfer (Lepidoptera, Nymphalidae, Danainae) from Taiwan. Transactions of the Lepidopterological Society of Japan 46 (4): 206-208.
54 Yen SH, Jean JL (1995) Life histories of Lethe gemina zaitha Fruhstorfer, 1914 (Lepidoptera: Nymphalidae, Satyridae). Transactions of the Lepidopterological Society of Japan 46 (3): 113-120.
55 Yen SH, Jean JL (1995) Notes on the life history of Sibataniozephyrus kuafui Hsu and Lin, 1994 (Lepidoptera: Lycaenidae). Chinese Journal of Entomology 15: 161-169.
56 Yen SH (1995) Notes on Iontha acerces Prout (Noctuidae, Catocalinae) from Taiwan. Japanese Heterocerists’ Journal 183: 126-128.
57 Yen SH (1995) Chrysartona stipata (Walker, 1854), a new addition to Zygaenidae (Procridinae) of Taiwan. Japanese Heterocerists’ Journal 182: 99-102.
58 Yang YP, Yen SH (1994) Deinostema in Taiwan (Scrophulariaceae). Botanical Bulletin of Academia Sinica (1994) 35: 61-63.
59 Yang YP, Yen SH, Huang S (1987) Two new additions of aquatic plants of Taiwan: Potamogeton maackianus and Utricularia minor. Botanical Bulletin of Academia Sinica (1987) 28: 49-53.

Monographic publications and book chapters

1 Yen SH (2003) Aquatic and semi-aquatic moths, with special reference to the Nymphulinae (Lepidoptera, Crambidae), in Y. Cathy (ed.), Aquatic Invertebrates of the Malaysian Region, Monash University and The Malaysian Academy of Sciences, Kuala Lumpur.
2 Yen SH, Yang PS (2002) Guide Book to the Insects Protected by the CITES Convention and the Wildlife Conservation Law in Taiwan (English version). Council of Agriculture, Executive Yuan, Taipei, 179 pp.
3 Yang YP, Yen SH, Lin CK (2002) Aquatic Flora of Taiwan. Council of Agriculture, Executive Yuan, Taiwan, R.O.C., Taipei, 378 pp.
4 Yen SH, Yang PS (2001) Guide Book of Plants in Campuses. Bureau of Environmental Protection, The Executive Yuan, Taipei, 106 pp.
5 Yen SH (2000) Lepidoptera, in Wu, W.J. (ed.), Illustration of pest insects. Bureau of animal and plant health inspection and quarantine, Council of Agriculture, Executive Yuan, Taipei, 156 pp.
6 Yen SH, Yang PS (2000) Illustrated Guide to Insects Protected by the CITES Convention and Wildlife Conservation Law in Taiwan. Council of Agriculture, Executive Yuen, R.O.C., Taipei, 112 pp.
7 Yang PS, Li CL, Yen SH, Chang ML (1999) Insects of the Kengting National Park-Guide book series of the Kengting National Park 11. Kengting National Park Agency, Hengchun, 120 pp.
8 Yen SH (1994) Aquatic and Wetland Plants in Keelung River Basin, pp.22-25, in P.S. Yang (ed.), Special Issue for the 100th anniversary of Taipei City, Taipei City Government, 63 pp.
9 Kuo CM, Chang YC, Yen SH (1988) Native Plants of Taiwan. Environmental Protection Education Committee, Taipei, 119 pp.

Links to some research projects I am involved in


Digital Archive Project for the Type Specimens and Literature of Taiwan Macrolepidoptera
2 Digital Archive Project for the Lepidopteran and Hostplant Specimens Housed in NSYSU
3 LepTree Project
4 Globiz
5 The Lepidoptera Taxome Project

Blogs of my lab

1 Care of Magical Creatures
2 LepLab
3 Digest of Taiwan Lepidopterology
4 Digileps News
5 Natural History of Takaw

Blogs of selected courses

1 General Biology (evolution, ecology & biodiversity parts)
2 Introduction to the Research Methods of Biological Sciences
3 Seminar
4 Systematic Biology & Practices of Biological Systematics
5 Mimicry Biology
6 Entomology
7 Scientific Illustration & Expression Skill